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More recent work by Devadoss and colleagues went beyond micronuclei alone, cytologically comparing buccal cells from nonsmokers, tobacco smokers, and tobacco chewers to evaluate for differences in their markers of genetic mutations, including karyorrhexis, karyolysis, pyknosis, binucleation, condensed nuclei, hyperchromatism, prominent nucleoli, broken egg nuclei, nuclear-cytoplasmic ratio, and irregular nuclear borders

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